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. 2018 Aug 21;8:12517. doi: 10.1038/s41598-018-31035-y

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© The Author(s) 2018

Open Access This article is licensed under a Creative Commons Attribution 4.0 International License, which permits use, sharing, adaptation, distribution and reproduction in any medium or format, as long as you give appropriate credit to the original author(s) and the source, provide a link to the Creative Commons license, and indicate if changes were made. The images or other third party material in this article are included in the article’s Creative Commons license, unless indicated otherwise in a credit line to the material. If material is not included in the article’s Creative Commons license and your intended use is not permitted by statutory regulation or exceeds the permitted use, you will need to obtain permission directly from the copyright holder. To view a copy of this license, visit http://creativecommons.org/licenses/by/4.0/.

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Non-linear viscoelastic properties of hippocampus obtained with dynamic indentation-controlled testing: frequency sweep (a) and oscillatory ramp (b). (a) Storage and loss moduli E′ and E″ increase over a frequency range of 1–10 Hz (data averaged over hippocampus; measured at ~7% strain, 0.2 μm oscillation amplitude; note the logarithmic scale on the x-axis). (b) E′ and E″ increase over the strain range of 4–10% (measured at 5.62 Hz oscillation frequency, 0.2 μm oscillation amplitude). The age of the animals is specified in the legend. Shadowed zones indicate the standard error of the mean, whereas R indicates the radius of the indenting tip and n the number of measurement points.