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. 2018 Jul 19;5(3):86–97. doi: 10.1159/000490491

Table 3.

Comparison of apoptosis with keratinization (modified from [1])

Apoptosis Keratinization
Removal of dead cells phagocytosis Keratinization desquamation
Cell membrane intact/PS exposure intact
Cell organelles encapsulated lysed
Nucleus fragmentation lost
 Chromatin condensation (+) (−)
 DNA fragmentation internucleosomal (ladder) lost
 TUNEL/ISTR (+) (−)
Nuclease caspase-activated DNase DNase 1L2/DNase 2
Cytoskeletal proteins cleaved (CK18/vimentin/actin) crosslinked → degraded
Involved enzymes caspase(-3) transglutaminase, caspase-14, KLK

PS, phosphatidylserine; TUNEL, TdT-mediated uptake of nick end labeling; ISTR, in situ 3′-tailing reaction; CK18, cytokeratin 18; KLK, kallikrein-kinin family.