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. 2018 Sep 3;8:13123. doi: 10.1038/s41598-018-31370-0

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© The Author(s) 2018

Open Access This article is licensed under a Creative Commons Attribution 4.0 International License, which permits use, sharing, adaptation, distribution and reproduction in any medium or format, as long as you give appropriate credit to the original author(s) and the source, provide a link to the Creative Commons license, and indicate if changes were made. The images or other third party material in this article are included in the article’s Creative Commons license, unless indicated otherwise in a credit line to the material. If material is not included in the article’s Creative Commons license and your intended use is not permitted by statutory regulation or exceeds the permitted use, you will need to obtain permission directly from the copyright holder. To view a copy of this license, visit http://creativecommons.org/licenses/by/4.0/.

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Hydrogen bonding pattern in the proton path. (a) Hydrogen bonding network between SB and Arg82. The 2F_obs − F_calc electron density map is shown at contour levels of 3σ (gray) and 5σ (dark gray). (b) Hydrogen bonding geometry around Tyr185 and Asp212. (c) The F_obs − F_calc omit map of water molecules in the proton path is shown at contour levels of 3σ (gray) and 5σ (magenta).