Figure 4.

Wooden (a,c) and stone anvils (b,d) used by capuchins to process structurally protected foods. Note accumulation of processed materials including old endocarps, coconut husks and shells in (a,b). (c) Hermit crab exoskeleton and shell remains processed with a stone tool on dead tree branch. (b,d) Stone hammer on stone anvil with T. catappa endocarps and halloween crab remains. The leaf stuck under the hammer in (d) indicates that the food was processed shortly before discovery. Percussion impacts can be observed on the crab limb.