Skip to main content
. 2018 Sep 4;6:e5436. doi: 10.7717/peerj.5436

Table 2. Behavioural ethogram of the study.

The behaviour list has been prepared based on preliminary observation of the flock and on previous studies on the Northern bald ibis as well as on relative species (Hirsch, 1979; Kopij, 1998; Pegoraro & Föger, 2001; Mark, 2007; Vargas-Ashby & Pankhurst, 2007; Clark et al., 2012; Fernández-Juricic, 2012; Moulton et al., 2013; Frigerio et al., 2016).

Individual behaviour  
  Alertness Moving the head, stretching the neck listening or watching something in the surrounding.
Attentive behaviour Being vigilant, scanning the environment.
  Comfort behaviour Tidying and cleaning the own feathers with the bill, scratching, bathing and sunbathing (fanning the wings to warm up in the sun).
  Exploration Looking for food probing the ground, tree branches, crevices and other elements of the aviary with the bill.
  Flight Flying or hopping from one point to the other of the enclosure.
  Walking Walking around the aviary.
  Maintenance Eating the food provided in the feeding point, defecating, drinking.
  Manipulation/play Manipulating food, objects, twigs, rocks or other inanimate items found in the aviary using the bill or the legs.
  Resting Standing on one or both legs, with the head turned back and tucked beneath the wings.
Social behaviour  
  Social affiliative behaviour  
Preening Tidying and cleaning the feathers of a conspecific with the bill.
  Other affiliative Greeting displays including head tossing, head rubbing, mutual bill shaking; observing conspecifics. 
  Social agonistic behaviour  
  Aggression Pecking towards approaching birds, hitting a conspecific with the bill or with the legs.
  Agonistic display Bill gaping, ruffling the helmsman feathers, moving or lunging toward conspecifics, touching slightly with the tip of the bill. Anti-predatory behaviour (e.g.: mobbing).
Not visible  
Out of sight The ibis spends time “out of sight” and it is not possible to see what the individual is doing. In particular, the bird is in sheltered areas such as nest boxes, rock crevices or behind trees and bushes.