Table 1.

Examples of Descriptors Extracted from Each Segment and Subsegment of an Image

Segment	Descriptors
Cytoplasm	Mean optical density, RGB
	Area
	Perimeter
	Contour
	Fourier decomposition
	Geometric primitive best fit
	Markov texture, r ε [1, 5], RGB
	Statistical moments, RGB
Nucleus	Mean optical density, RGB
	Area
	Perimeter
	Ratio of nuclear area to cytoplasm area
	Eccentricity
	Contour
	Fourier decomposition
	Geometric primitive best fit
	Markov texture, r ε [1, 5], RGB
	Statistical moments, RGB
Nuclear subsegments	Markov texture, r = 1, RGB
	Statistical moments, RGB
Nucleolus	Mean optical density, RGB
	Area
	Perimeter
	Nucleolus/nucleus area ratio
	Eccentricity
	Contour
	Fourier decomposition
	Geometric primitive best fit
	Markov texture, r = 1, RGB
	Statistical moments, RGB
Note: r values indicate the window size of the gray-level co-occurrence matrix expressed in horizontal pixels.