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. 2018 Sep 26;8:14415. doi: 10.1038/s41598-018-32686-7

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© The Author(s) 2018

Open Access This article is licensed under a Creative Commons Attribution 4.0 International License, which permits use, sharing, adaptation, distribution and reproduction in any medium or format, as long as you give appropriate credit to the original author(s) and the source, provide a link to the Creative Commons license, and indicate if changes were made. The images or other third party material in this article are included in the article’s Creative Commons license, unless indicated otherwise in a credit line to the material. If material is not included in the article’s Creative Commons license and your intended use is not permitted by statutory regulation or exceeds the permitted use, you will need to obtain permission directly from the copyright holder. To view a copy of this license, visit http://creativecommons.org/licenses/by/4.0/.

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(A) Model of bacterium showing wrapped-wrapped swimming with both bundles left-handed while both motors rotate CCW with the same torque, set at 2000 pN nm. (B) Swimming speed and body rotation rate as a function of undetermined rolling rotation rates of each bundle. (C) Rigid-body rotation rates of each bundle as a function of undetermined rolling rotation rates. (D) Model of wrapped-wrapped swimming with one bundle left-handed and the other bundle right-handed while both motors rotate CCW with same torque set at 2000 pN nm. (E) Swimming speed and body rotation rate as a function of undetermined rolling rotation rates of each bundle. (F) Rigid-body rotation rates of each bundle as a function of undetermined rolling rotation rates.