Fig 3. The full model is capable of performing a range of conditioning tasks.

With modification of only the experimental protocol, our full model can successfully perform a range of conditioning tasks which can be performed by restrained (using the Proboscis Extention Reflex (PER) paradigm) and free flying bees. Performance closely matches experimental data with real bees (e.g. A: [46], B: [47], C & D: [48]).