Net diversification rates are similar among warm marine regions, and highest in the cold oceans. (a) The relationship between weighted mean net diversification rates for each region, and the regions' total richness. Means for each region are the mean net diversification rates of families or genera present, weighted by each clade's richness in that region. Here, rates were calculated using the crown age and assuming ɛ = 0.5. Richness was log10-transformed prior to performing regressions. (b) Net diversification rates inferred by best-fit HiSSE models. We performed three binary comparisons by combining regions (Methods) and pruning the phylogeny to include only species in the regions of interest. For the analyses shown, only species restricted to the Indo-West Pacific (IWP), cold oceans, and freshwater habitats were assigned to each of these groups. Alternative HiSSE analyses assigned widespread species to these categories. Rates were averaged for each region across hidden states, based on the distribution of the hidden states among regions. Details of alternative analyses, hidden states and GeoHiSSE are given in the electronic supplementary material, appendix S3.