(A-B) Final states for simulations of chronic TCDD exposure. Scenarios
were parameterized via AhR-mediated fold-changes in EGFR (as indicated from 1.1x
to 1.3x) for (A) the low-hysteresis version of the EGF-TGFβ3 switch
(n=54, tipping point ~1.2x) and (B) the high-hysteresis version (n=34,
tipping point ~1.2x). Insets show corresponding EGF and TGFβ3
signal gradients. The phenotype of a thickened MES preventing mesenchymal
confluence is similar in appearance to the histology of palatal shelves from a
TGFβ3 knockout (with or without additional Alk mutations)44. (C-D) Time-lapse images from
simulations of transient acute exposure to ATRA parameterized as time-dependent
EGFR fold-changes (top). Palate fusion was delayed using the low-hysteresis
switch (n=24, tipping point >1.8x) (C), but failed using the
high-hysteresis switch (n=16, tipping point 1.5x) (D) despite being subjected to
a smaller maximum EGFR fold-change. Complete time-lapse image sets are available
for specific examples as Supplemental Movies S3 (panel A, 1.15x), S4 (panel B, 1.2x), S5 (panel C), and S6 (panel D).