Table II.
Gene | SNP | Nucleotide substitutiona | Amino acid substitution | Chromosome | Position | MAF, % | Allele OR | P-value, allele frequency | FDR, allele frequency |
---|---|---|---|---|---|---|---|---|---|
PLCB2 | rs200787930 | C/T | E1106K | 15 | 40289298 | 1.2 | 0.03 | 1.24×10−29 | 1.56×10−26 |
MARCH1 | rs61734696 | G/T | Q137K | 4 | 164197303 | 1.2 | 0.03 | 2.09×10−29 | 2.54×10−26 |
VPS33B | rs199921354 | C/T | R80Q | 15 | 91013841 | 1.2 | 0.03 | 2.76×10−29 | 3.30×10−26 |
CXCL8 | rs188378669 | G/T | E31* | 4 | 73741568 | 1.2 | 0.03 | 3.15×10−29 | 3.70×10−26 |
TMOD4 | rs115287176 | G/A | R277W | 1 | 151170961 | 1.2 | 0.03 | 1.21×10−28 | 1.39×10−25 |
COL6A3 | rs146092501 | C/T | E1386K | 2 | 237371861 | 1.2 | 0.04 | 2.93×10−28 | 3.27×10−25 |
ZNF77 | rs146879198 | G/A | R340* | 19 | 2934109 | 1.2 | 0.04 | 2.92×10−28 | 3.27×10−25 |
ADGRL3 | rs192210727 | G/T | R580I | 4 | 61909615 | 1.3 | 0.10 | 2.92×10−23 | 3.06×10−20 |
OR52E4 | rs11823828 | T/G | F227L | 11 | 5884973 | 36.6 | 1.54 | 3.40×10−21 | 3.35×10−18 |
ALDH2 | rs671 | G/A | E504K | 12 | 111803962 | 27.6 | 1.41 | 4.12×10−15 | 3.78×10−12 |
ACAD10 | rs11066015 | G/A | 12 | 111730205 | 27.5 | 1.41 | 4.92×10−15 | 4.45×10−12 | |
BRAP | rs3782886 | A/G | 12 | 111672685 | 29.3 | 1.37 | 4.38×10−13 | 3.71×10−10 | |
HECTD4 | rs11066280 | T/A | 12 | 112379979 | 29.0 | 1.37 | 6.94×10−13 | 5.73×10−10 | |
HECTD4 | rs2074356 | C/T | 12 | 112207597 | 25.4 | 1.36 | 1.21×10−11 | 9.78×10−9 | |
NAA25 | rs12231744 | C/T | R876K | 12 | 112039251 | 35.1 | 0.77 | 1.68×10−9 | 1.24×10−6 |
GOSR2 | rs1052586 | T/C | 17 | 46941097 | 48.7 | 0.79 | 3.94×10−8 | 2.61×10−5 | |
ATXN2 | rs7969300 | T/C | N248S | 12 | 111555908 | 38.8 | 0.79 | 4.41×10−8 | 2.87×10−5 |
LILRB2 | rs73055442 | C/T | R103H | 19 | 54279838 | 1.6 | 44.10 | 2.00×10−7 | 1.20×10−4 |
rs12229654 | T/G | 12 | 110976657 | 22.5 | 1.28 | 2.09×10−7 | 1.24×10−4 | ||
LOC107987429 | rs2844533 | T/C | 6 | 31383025 | 15.3 | 1.32 | 3.49×10−7 | 1.95×10−4 | |
MTFR2 | rs143974258 | G/A | R360* | 6 | 136231355 | 3.3 | 0.05 | 6.66×10−7 | 3.60×10−4 |
PSORS1C1 | rs3130559 | C/T | 6 | 31129524 | 44.2 | 0.82 | 1.51×10−6 | 7.74×10−4 | |
rs2596548 | G/T | 6 | 31362769 | 5.4 | 1.51 | 1.83×10−6 | 9.21×10−4 | ||
EIF3L | rs9466 | T/C | 22 | 37877742 | 21.6 | 1.28 | 1.96×10−6 | 9.77×10−4 | |
LPGAT1 | rs150552771 | T/C | K200E | 1 | 211783358 | 5.0 | 7.14 | 2.26×10−6 | 0.0011 |
LAIR2 | rs34429135 | T/A | F115Y | 19 | 54508164 | 2.5 | ND | 2.70×10−6 | 0.0013 |
rs2523644 | A/G | 6 | 31374707 | 8.1 | 1.40 | 2.75×10−6 | 0.0013 | ||
rs10757278 | A/G | 9 | 22124478 | 49.5 | 0.83 | 2.92×10−6 | 0.0014 | ||
CCHCR1 | rs130067 | T/G | E328D | 6 | 31150734 | 33.2 | 0.81 | 3.10×10−6 | 0.0015 |
TCHP | rs74416240 | G/A | 12 | 109904793 | 13.3 | 1.30 | 3.25×10−6 | 0.0015 | |
rs1333049 | G/C | 9 | 22125504 | 49.4 | 1.20 | 3.95×10−6 | 0.0018 | ||
CDKN2B-AS1 | rs4977574 | A/G | 9 | 22098575 | 47.1 | 1.21 | 4.18×10−6 | 0.0019 | |
CDKN2B-AS1 | rs2383207 | G/A | 9 | 22115960 | 33.7 | 0.81 | 4.86×10−6 | 0.0022 | |
SLC16A1 | rs1049434 | T/A | D490E | 1 | 112913924 | 34.7 | 0.82 | 5.76×10−6 | 0.0025 |
GIT2 | rs925368 | T/C | N389S | 12 | 109953174 | 12.5 | 1.30 | 6.02×10−6 | 0.0026 |
rs1333048 | A/C | 9 | 22125348 | 49.6 | 1.20 | 6.46×10−6 | 0.0028 | ||
rs2523578 | T/C | 6 | 31360765 | 8.1 | 1.39 | 6.54×10−6 | 0.0028 | ||
rs404890 | G/T | 6 | 32231090 | 30.5 | 1.22 | 8.90×10−6 | 0.0037 | ||
APOE | rs7412 | C/T | R176C | 19 | 44908822 | 4.3 | 0.60 | 1.06×10−5 | 0.0043 |
CCHCR1 | rs130071 | G/A | 6 | 31148433 | 5.1 | 1.52 | 1.06×10−5 | 0.0043 | |
rs602633 | C/A | 1 | 109278889 | 7.6 | 0.69 | 1.15×10−5 | 0.0046 | ||
CELSR2 | rs12740374 | G/T | 1 | 109274968 | 7.7 | 0.69 | 1.15×10−5 | 0.0046 | |
MKI67 | rs145121731 | G/A | S2722L | 10 | 128102595 | 1.5 | 2.04 | 1.20×10−5 | 0.0047 |
CUBN | rs78201384 | C/T | E304K | 10 | 17111024 | 2.7 | 0.52 | 1.38×10−5 | 0.0054 |
PSORS1C3 | rs887466 | T/C | 6 | 31175734 | 41.1 | 1.20 | 1.38×10−5 | 0.0054 | |
PSORS1C1 | rs3094663 | G/A | 6 | 31139310 | 30.9 | 1.20 | 1.40×10−5 | 0.0054 | |
rs10853110 | A/G | 17 | 49241052 | 39.2 | 1.20 | 1.49×10−5 | 0.0057 | ||
WDR37 | rs10794720 | C/T | 10 | 1110225 | 8.5 | 0.71 | 1.52×10−5 | 0.0057 | |
CELSR2 | rs629301 | A/C | 1 | 109275684 | 7.8 | 0.70 | 1.52×10−5 | 0.0057 | |
SKIV2L | rs592229 | G/T | 6 | 31962664 | 42.4 | 1.20 | 1.57×10−5 | 0.0058 | |
rs12182351 | T/C | 6 | 32233930 | 29.8 | 1.22 | 1.59×10−5 | 0.0059 | ||
POU5F1 | rs3130503 | G/A | 6 | 31169388 | 29.5 | 1.20 | 1.64×10−5 | 0.0060 | |
PSORS1C3 | rs1265155 | T/C | 6 | 31175917 | 41.1 | 1.19 | 1.68×10−5 | 0.0061 | |
CELSR2 | rs646776 | A/G | 1 | 109275908 | 7.7 | 0.70 | 1.70×10−5 | 0.0062 | |
rs2596503 | C/T | 6 | 31353033 | 19.3 | 1.24 | 1.75×10−5 | 0.0063 | ||
TRPM1 | rs2241493 | T/C | N54S | 15 | 31070149 | 12.6 | 0.76 | 1.81×10−5 | 0.0065 |
CCDC141 | rs13419085 | T/C | N1170S | 2 | 178837710 | 1.8 | 0.46 | 1.92×10−5 | 0.0068 |
VARS2 | rs9394021 | A/G | Q777R | 6 | 30925350 | 44.9 | 0.84 | 1.98×10−5 | 0.0069 |
SFTA2 | rs2286655 | T/C | 6 | 30931969 | 44.9 | 1.19 | 1.99×10−5 | 0.0069 | |
rs3873334 | T/C | 6 | 30928370 | 44.9 | 1.19 | 1.98×10−5 | 0.0069 | ||
rs9261800 | C/G | 6 | 30408822 | 2.8 | 7.21 | 2.02×10−5 | 0.0069 | ||
TCF19 | rs3130453 | C/T | 6 | 31157072 | 34.4 | 0.83 | 2.10×10−5 | 0.0072 | |
C21orf59 | rs76974938 | C/T | D67N | 21 | 32609946 | 2.4 | 0.00 | 2.14×10−5 | 0.0073 |
DDR1 | rs2239518 | T/C | 6 | 30897948 | 44.9 | 1.19 | 2.19×10−5 | 0.0074 | |
CDSN | rs3130984 | C/T | S143N | 6 | 31117187 | 13.4 | 1.29 | 2.20×10−5 | 0.0074 |
rs197932 | T/C | 17 | 46896981 | 26.9 | 0.82 | 2.23×10−5 | 0.0075 | ||
CDSN | rs3130981 | C/T | D527N | 6 | 31116036 | 13.6 | 1.29 | 2.30×10−5 | 0.0075 |
MICB-DT | rs3132469 | C/T | 6 | 31488790 | 5.3 | 1.46 | 2.41×10−5 | 0.0078 | |
HLA-DQB1 | rs1049056 | C/A | A6S | 6 | 32666592 | 11.9 | 1.30 | 2.51×10−5 | 0.0081 |
DDR1 | rs2239517 | A/G | 6 | 30897338 | 44.6 | 1.19 | 2.59×10−5 | 0.0083 | |
CCHCR1 | rs1265110 | G/A | 6 | 31151645 | 30.2 | 0.83 | 2.69×10−5 | 0.0085 | |
CCDC63 | rs10774610 | T/C | 12 | 110902439 | 23.7 | 1.22 | 2.76×10−5 | 0.0087 | |
GTF2H4 | rs2284176 | C/T | 6 | 30907845 | 44.6 | 1.19 | 2.80×10−5 | 0.0088 | |
GTF2H4 | rs3909130 | G/A | 6 | 30906388 | 44.6 | 1.19 | 2.84×10−5 | 0.0089 | |
GTF2H4 | rs916920 | G/A | 6 | 30909425 | 44.7 | 1.19 | 2.85×10−5 | 0.0089 | |
rs1264569 | A/G | 6 | 30397543 | 4.6 | 1.49 | 2.98×10−5 | 0.0092 | ||
CACNA1D | rs35874056 | G/A | G460S | 3 | 53702798 | 2.0 | 25.00 | 3.09×10−5 | 0.0094 |
rs9468845 | A/G | 6 | 30901816 | 44.7 | 1.19 | 3.12×10−5 | 0.0094 | ||
DDR1 | rs8408 | C/T | 6 | 30899889 | 44.7 | 1.19 | 3.11×10−5 | 0.0094 | |
DDR1 | rs7756521 | C/T | 6 | 30880476 | 44.7 | 1.19 | 3.10×10−5 | 0.0094 | |
CDKN2B-AS1 | rs1011970 | G/T | 9 | 22062135 | 5.6 | 1.41 | 3.15×10−5 | 0.0095 | |
ADAT1 | rs145161932 | T/C | R57G | 16 | 75612670 | 1.4 | 0.39 | 3.29×10−5 | 0.0098 |
POU5F1 | rs885950 | T/G | 6 | 31172375 | 34.0 | 0.83 | 3.28×10−5 | 0.0098 | |
DDR1 | rs4618569 | A/G | 6 | 30887474 | 44.7 | 1.19 | 3.41×10−5 | 0.0101 | |
KRT13 | rs146918776 | A/G | Y281H | 17 | 41502993 | 1.5 | 1.94 | 3.51×10−5 | 0.0103 |
rs2523638 | G/A | 6 | 31376496 | 43.1 | 1.19 | 3.53×10−5 | 0.0103 | ||
PSRC1 | rs599839 | A/G | 1 | 109279544 | 7.9 | 0.71 | 3.52×10−5 | 0.0103 | |
rs9275141 | G/T | 6 | 32683340 | 26.4 | 1.21 | 3.62×10−5 | 0.0105 | ||
CCDC63 | rs10849915 | T/C | 12 | 110895818 | 23.6 | 1.22 | 3.63×10−5 | 0.0105 | |
HLA-DRA | rs3177928 | G/A | 6 | 32444658 | 5.9 | 1.41 | 3.81×10−5 | 0.0108 | |
OAS3 | rs2072134 | C/T | 12 | 112971371 | 17.6 | 1.24 | 4.06×10−5 | 0.0114 | |
USP45 | rs41288947 | C/G | T521R | 6 | 99446210 | 14.9 | 1.26 | 4.11×10−5 | 0.0115 |
CCHCR1 | rs1265109 | A/C | 6 | 31151812 | 48.2 | 1.18 | 4.16×10−5 | 0.0116 | |
LOC101929163 | rs6930777 | C/T | 6 | 32383789 | 5.5 | 1.43 | 4.45×10−5 | 0.0122 | |
rs7333181 | G/A | 13 | 111568950 | 2.5 | 0.54 | 4.45×10−5 | 0.0122 | ||
DDR1 | rs1264323 | T/C | 6 | 30888130 | 38.8 | 1.19 | 4.48×10−5 | 0.0122 | |
LINC00243 | rs3094111 | G/A | 6 | 30820414 | 14.7 | 1.25 | 4.52×10−5 | 0.0123 | |
rs10484561 | T/G | 6 | 32697643 | 5.9 | 1.41 | 4.55×10−5 | 0.0123 | ||
PSORS1C1 | rs3130558 | G/C | 6 | 31129406 | 13.7 | 1.27 | 4.59×10−5 | 0.0124 | |
HLA-DQB1 | rs1049060 | T/A | S27T | 6 | 32666529 | 28.8 | 1.20 | 4.92×10−5 | 0.0131 |
rs2844650 | G/A | 6 | 30934756 | 4.7 | 1.47 | 4.99×10−5 | 0.0131 | ||
DDR1 | rs3132572 | T/C | 6 | 30893952 | 4.7 | 1.47 | 4.99×10−5 | 0.0131 | |
CCHCR1 | rs1265115 | T/G | 6 | 31149298 | 47.7 | 1.18 | 4.96×10−5 | 0.0131 | |
CCHCR1 | rs3094225 | T/C | 6 | 31145275 | 48.4 | 1.18 | 4.93×10−5 | 0.0131 | |
LOC107987453 | rs3129987 | C/T | 6 | 30798427 | 14.5 | 1.25 | 5.05×10−5 | 0.0132 | |
DPCR1 | rs2517451 | A/G | 6 | 30946974 | 4.7 | 1.47 | 5.11×10−5 | 0.0133 | |
KIAA1551 | rs10771894 | A/G | S352G | 12 | 31982009 | 32.4 | 1.19 | 5.18×10−5 | 0.0134 |
rs13427905 | C/T | 2 | 71846585 | 18.5 | 0.80 | 5.22×10−5 | 0.0134 | ||
ABCA1 | rs1883025 | G/A | 9 | 104902020 | 28.8 | 0.83 | 5.46×10−5 | 0.0139 | |
SFTA2 | rs2253705 | G/A | 6 | 30932317 | 18.0 | 1.23 | 5.60×10−5 | 0.0141 | |
PLUT | rs954750 | G/A | 13 | 27889801 | 48.3 | 1.18 | 5.86×10−5 | 0.0146 | |
TCF19 | rs1419881 | T/C | 6 | 31162816 | 48.1 | 1.18 | 6.37×10−5 | 0.0156 | |
rs13209234 | G/A | 6 | 32448198 | 5.9 | 1.41 | 6.47×10−5 | 0.0158 | ||
PSORS1C1 | rs1265100 | T/C | 6 | 31137533 | 32.2 | 0.83 | 6.55×10−5 | 0.0159 | |
YEATS2 | rs76174573 | G/T | C1232F | 3 | 183804099 | 3.7 | 0.61 | 6.74×10−5 | 0.0162 |
ABO | rs1053878 | C/T | P156L | 9 | 133256264 | 22.8 | 1.20 | 6.78×10−5 | 0.0162 |
rs4014195 | C/G | 11 | 65739351 | 16.6 | 1.24 | 6.78×10−5 | 0.0162 | ||
SFTA2 | s2253588 | C/G | 6 | 30931600 | 23.6 | 1.21 | 6.93×10−5 | 0.0165 | |
CYP4F8 | rs201166643 | C/A | R488S | 19 | 15629257 | 1.1 | ND | 7.00×10−5 | 0.0165 |
NAXE | rs7516274 | C/G | L19V | 1 | 156591859 | 1.8 | 0.48 | 7.18×10−5 | 0.0169 |
rs10757283 | T/C | 9 | 22134173 | 33.8 | 0.84 | 7.25×10−5 | 0.0170 | ||
BTNL2 | rs28362680 | G/A | A202V | 6 | 32403039 | 39.7 | 0.85 | 7.40×10−5 | 0.0171 |
BTNL2 | rs10947262 | C/T | 6 | 32405535 | 39.7 | 0.85 | 7.40×10−5 | 0.0171 | |
KRT27 | rs17558532 | C/T | A284T | 17 | 40779624 | 3.6 | 0.62 | 7.71×10−5 | 0.0176 |
GTF2H4 | rs3130780 | G/T | 6 | 30906531 | 18.0 | 1.23 | 7.71×10−5 | 0.0176 | |
rs2532934 | T/C | 6 | 30926982 | 24.1 | 1.20 | 7.74×10−5 | 0.0176 | ||
VARS2 | rs753725 | G/A | 6 | 30923094 | 24.1 | 1.20 | 7.68×10−5 | 0.0176 | |
PLUT | rs11619319 | A/G | 13 | 27913462 | 48.1 | 1.18 | 7.64×10−5 | 0.0176 | |
rs3095273 | C/T | 6 | 29598592 | 5.5 | 1.41 | 8.16×10−5 | 0.0184 | ||
TNS1 | rs918949 | C/T | V1590I | 2 | 217809974 | 42.8 | 0.85 | 8.39×10−5 | 0.0188 |
LINC00243 | rs3130785 | C/T | 6 | 30828961 | 14.6 | 1.24 | 8.37×10−5 | 0.0188 | |
VARS2 | rs2249464 | C/T | R309W | 6 | 30920384 | 24.1 | 1.20 | 9.39×10−5 | 0.0207 |
rs3095345 | A/G | 6 | 30854636 | 17.9 | 1.22 | 9.37×10−5 | 0.0207 | ||
ITGB8 | rs80015015 | G/A | C481Y | 7 | 20401881 | 7.1 | 1.35 | 1.01×10−4 | 0.0220 |
VARS2 | rs885905 | C/T | 6 | 30922654 | 23.4 | 1.20 | 1.07×10−4 | 0.0232 | |
LIPE | rs34052647 | G/A | R611C | 19 | 42407617 | 5.5 | 1.39 | 1.16×10−4 | 0.0249 |
PHACTR1 | rs9369640 | A/C | 6 | 12901209 | 9.1 | 0.74 | 1.30×10−4 | 0.0275 | |
BTNL2 | rs41417449 | T/C | M295V | 6 | 32396234 | 23.0 | 0.83 | 1.35×10−4 | 0.0280 |
BTNL2 | rs41441651 | C/T | D336N | 6 | 32396111 | 23.0 | 0.83 | 1.35×10−4 | 0.0280 |
BTNL2 | rs28362675 | C/A | E454* | 6 | 32394744 | 23.0 | 0.83 | 1.35×10−4 | 0.0280 |
BTNL2 | rs78587369 | G/A | T165I | 6 | 32403150 | 23.0 | 0.83 | 1.35×10−4 | 0.0280 |
BTNL2 | rs3763315 | G/T | 6 | 32408877 | 23.0 | 0.83 | 1.35×10−4 | 0.0280 | |
BTNL2 | rs2076528 | T/G | 6 | 32396417 | 23.0 | 0.83 | 1.35×10−4 | 0.0280 | |
PRKG1 | rs9414827 | G/A | 10 | 51137314 | 10.1 | 0.76 | 1.37×10−4 | 0.0282 | |
rs6537384 | T/G | 4 | 145949613 | 28.8 | 1.19 | 1.43×10−4 | 0.0294 | ||
rs6067640 | G/A | 20 | 51092837 | 38.5 | 0.85 | 1.48×10−4 | 0.0302 | ||
rs10514995 | A/G | 5 | 66443611 | 48.7 | 1.16 | 1.51×10−4 | 0.0306 | ||
BTNL2 | rs34423804 | T/A | V283D | 6 | 32396269 | 23.0 | 0.83 | 1.63×10−4 | 0.0329 |
PHACTR1 | rs9349379 | G/A | 6 | 12903725 | 34.2 | 0.85 | 1.69×10−4 | 0.0341 | |
STIM1 | rs116855870 | A/G | 11 | 4055527 | 1.1 | 1.93 | 1.71×10−4 | 0.0343 | |
ZNF142 | rs3821033 | C/T | A1313T | 2 | 218642579 | 11.2 | 1.26 | 1.78×10−4 | 0.0355 |
LINC00354 | rs4907518 | G/A | 13 | 111898209 | 45.6 | 0.85 | 1.82×10−4 | 0.0362 | |
TNS3 | rs11763932 | G/A | 7 | 47567880 | 42.0 | 0.85 | 1.91×10−4 | 0.0378 | |
BTRC | rs2270439 | C/A | P566H | 10 | 101550817 | 3.5 | 0.63 | 1.94×10−4 | 0.0381 |
MIA3 | rs2936051 | A/G | E881G | 1 | 222629862 | 40.1 | 0.85 | 1.96×10−4 | 0.0384 |
rs6825911 | C/T | 4 | 110460482 | 45.9 | 0.86 | 2.01×10−4 | 0.0391 | ||
VNN1 | rs2294757 | G/A | T26I | 6 | 132713959 | 37.4 | 0.85 | 2.02×10−4 | 0.0393 |
ZNF860 | rs140232911 | C/T | S161L | 3 | 31989561 | 10.4 | 0.44 | 2.09×10−4 | 0.0406 |
rs838880 | C/T | 12 | 124777047 | 47.5 | 1.16 | 2.23×10−4 | 0.0430 | ||
MIA3 | rs2936052 | A/G | K605R | 1 | 222629034 | 34.4 | 0.85 | 2.26×10−4 | 0.0430 |
DTNBP1 | rs2743868 | G/A | 6 | 15625577 | 31.6 | 1.18 | 2.26×10−4 | 0.0430 | |
MON2 | rs11174549 | A/G | I1385V | 12 | 62565357 | 5.0 | 1.40 | 2.26×10−4 | 0.0430 |
rs507666 | G/A | 9 | 136149399 | 27.8 | 1.18 | 2.26×10−4 | 0.0430 | ||
FAM170B | rs73302786 | G/T | D252E | 10 | 49131709 | 3.5 | 1.47 | 2.36×10−4 | 0.0445 |
PSORS1C3 | rs3131018 | G/T | 6 | 31175805 | 15.7 | 1.23 | 2.36×10−4 | 0.0445 | |
PIEZO2 | rs35033671 | C/A | C1148F | 18 | 10759842 | 11.0 | 1.27 | 2.39×10−4 | 0.0448 |
SLC22A3 | rs1810126 | C/T | 6 | 160451119 | 49.1 | 0.86 | 2.46×10−4 | 0.0460 | |
PANK1 | rs11185790 | G/A | 10 | 89612776 | 46.9 | 1.16 | 2.57×10−4 | 0.0481 | |
GFY | rs73053944 | C/G | T203S | 19 | 49427038 | 2.9 | 1.51 | 2.58×10−4 | 0.0481 |
RNF2 | rs1046592 | A/G | 1 | 185100429 | 33.9 | 0.85 | 2.63×10−4 | 0.0488 |
Allele frequencies were analyzed using Fisher's exact test.
Major allele/minor allele. SNP, single nucleotide polymorphisms; MAF, minor allele frequency; OR, odds ratio; FDR, false discovery rate; ND, not determined.