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. 2018 Oct 26;8:15839. doi: 10.1038/s41598-018-34134-y

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© The Author(s) 2018

Open Access This article is licensed under a Creative Commons Attribution 4.0 International License, which permits use, sharing, adaptation, distribution and reproduction in any medium or format, as long as you give appropriate credit to the original author(s) and the source, provide a link to the Creative Commons license, and indicate if changes were made. The images or other third party material in this article are included in the article’s Creative Commons license, unless indicated otherwise in a credit line to the material. If material is not included in the article’s Creative Commons license and your intended use is not permitted by statutory regulation or exceeds the permitted use, you will need to obtain permission directly from the copyright holder. To view a copy of this license, visit http://creativecommons.org/licenses/by/4.0/.

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Etruscan Tomba degli Scudi, Tarquinia, Italy. (a) Hand-drawn map of the tomb from Maria Cristina Tomassetti, based on measurements taken during the restoration. The walls of the Stanza degli Scudi are indicated with letters. (b) Picture of the northern wall, (c) southern wall, (d) western wall, (e) and eastern wall. The red stars mark the sampling sites for microbiological analysis, and the yellow stars indicate the metagenomics sampling sites. A fine and uniform dry white patina is present on the wall surfaces with a gradient from the western wall towards the central atrium. The eastern wall has less patina than the other walls. Photographer: Domenico Ventura.