Table 2:

Systems and linker lengths modeled. Unless otherwise stated, no linker histones are included. NFRs are denoted in brackets. For exact values of linker length pairs analyzed in systems 8 and 9, see figures.

	Designation	Composition Linker Length (bp)	Linker Length Sequence (bp)	Ref
1	Uniform	100% 18 (NRL = 165)	Uniform (18,18,18,18,18,18,18,18,18,18,...)	41
2	Uniform	100 % 27 (NRL = 174)	Uniform (27,27,27,27,27,27,27,27,27,27,...)	41
3	Life-like	70% 18, 20% 27, 10% 36	27 36 18 27 18 27 27 18 27 36 18 18 18 18 36 18 18 18 36 27 18 18 18 36 36 36 27 18 18 18 36 18 18 18 27 27 36 18 27 27 27 36 18 27 18 18 36 18 27 18 27 36 18 18 36 36 27 27 27 36 18 27 18 27 18 18 27 18 27 36 18 27 18 18 18 27 18 36 27 18 18 27 27 27 27 18 27 36 36 27 18 18 27 36 18 18 27 18 27 27	23
4	Life-like with NFRs Short NFRs: [72 bp] Long NFRs: [162 bp]	70% 18, 20% 27, 10% 36	[162] 36 18 27 18 27 27 18 [72] 36 18 18 18 18 36 18 18 [72] 36 27 18 18 18 36 [72] 36 27 18 18 [72] 36 18 18 18 27 27 36 [72] 27 27 27 36 18 27 [162] 18 36 18 27 18 27 36 18 18 36 36 27 [72] 27 [162] 18 27 18 27 18 18 27 18 27 36 18 27 [72] 18 18 27 18 36 27 18 18 27 27 27 27 18 27 36 36 27 18 18 27 36 18 [162] 27 18 27 27	15,23
5	Gene encoding-like Short NFRs: [72 bp] Long NFRs: [162 bp]	70% 18, 20% 27, 10% 36 with short linkers at the 5′ end	5′ [162] 18 18 18 18 18 27 36 [72] 18 18 18 27 18 36 27 27 [72] 18 18 18 36 18 27 [72] 18 18 18 27 [72] 18 18 18 18 18 27 36 [72] 18 18 36 27 18 27 [162] 18 18 18 27 18 36 27 18 27 18 27 36 [72] 18 [162] 18 18 18 27 18 18 18 18 27 36 18 27 [72] 18 18 18 18 18 27 18 18 27 18 36 27 18 18 36 36 18 18 18 27 36 18 [162] 18 18 18 18 3′	15,23
6	Gene encoding-like $\frac{1}{2}\text{LH}$	70% 18, 20% 27, 10% 36	″	15,23
7	Gene encoding-like +LH	70% 18, 20% 27, 10% 36	″	15,23
8	Random (7 pairs of linker lengths)	50% x (18–71), 50% y (18–71)	Random (x,y,y,x,x,y,y,x,y,x,y...)
9	Alternating (7 pairs of linker lengths)	50% x (18–71), 50% y (18–71)	Alternating (x,y,x,y,x,y,x,y,x,y...)	42