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Study 1: forced expiratory volume in 1 s (FEV1) changes caused by RPL554 (6 mg), salbutamol (200 µg) and ipratropium (40 µg) alone and in combination (single doses). a) Peak FEV1 and b) average FEV1(0–8 h). Data are presented as mean±sem. **: p<0.01.
