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Study 2: a) Residual volume (RV) and b) specific airway conductance (sGaw) changes on day 3 caused by tiotropium (18 µg), tiotropium (18 µg)+RPL554 (1.5 mg) and tiotropium (18 µg)+RPL554 (6 mg). The change from baseline (pre-dose on day 1) is shown. Data are presented as mean±sem. **: p<0.01.
