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. 2018 Nov 8;8:16590. doi: 10.1038/s41598-018-34989-1

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© The Author(s) 2018

Open Access This article is licensed under a Creative Commons Attribution 4.0 International License, which permits use, sharing, adaptation, distribution and reproduction in any medium or format, as long as you give appropriate credit to the original author(s) and the source, provide a link to the Creative Commons license, and indicate if changes were made. The images or other third party material in this article are included in the article’s Creative Commons license, unless indicated otherwise in a credit line to the material. If material is not included in the article’s Creative Commons license and your intended use is not permitted by statutory regulation or exceeds the permitted use, you will need to obtain permission directly from the copyright holder. To view a copy of this license, visit http://creativecommons.org/licenses/by/4.0/.

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The swimming plus-maze test (SPM): apparatus, behaviour and selection of variables. (a) 3D (top-left) and 2D drawings of apparati for larvae (bottom-left) and juveniles (bottom-right). Dashed lines indicate the boundary of areas (d: deep arm; s: shallow arm; c: center zone (intersection + ramps)). For exact dimensions of the apparati see Table 1. (b) Percentage of time spent in each zone of the SPM test; (c) representative heatmap of exploration (top) and fine resolution analysis of deep arm activity (bottom)(dashed lines indicate mean ± SEM). (d) Correlation between behavioural measures of the SPM test. Size and colour intensity of dots in the intersection of variable abbreviations are in linear association with the correlation coefficient between the two variables. Bigger dot indicate stronger association. Blue dots indicate positive, red ones indicate negative correlation. *Shows significant correlation between variables. DAE: deep/total arm entries, C: choice index, SAE: shallow/total arm entries, VELO: mean velocity, TAE: total arm entries.