| DCCs group-1 | DCCs group-2 | DCCs group-3 | |||
| Gene group-11 | +F11 | 0 | 0 | ||
| Gene group-12 | –F12 | 0 | 0 | ||
| Ytlq = | Gene group-21 | 0 | +F21 | 0 | +N(0, σ2) |
| Gene group-22 | 0 | –F22 | 0 | ||
| Gene group-3 | 0 | 0 | 0 | (2) |
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