Table 1.

MST network metrics

Symbol	Concept	Explanation
N	Nodes	Number of nodes in the MST
M	Links	Number of links in the MST
D	Degree	Number of neighbors for a given node in the MST
L	Leaf fraction	Fraction of leaf nodes in the MST where a leaf node is defined as a node with degree one
d	Diameter	Longest shortest path in the MST
Ecc	Eccentricity	Longest distance between a reference node and any other node
BC	Betweenness centrality	Fraction of all shortest paths that pass through a particular node
k	Kappa	Measure of the broadness of the degree distribution
Th	Tree hierarchy	A hierarchical metric that quantifies the trade-off between large scale integration in the MST and the overload of central nodes
R	Degree correlation	Correlation between the degrees of a node and the degree of the neighboring vertices to which it is connected