GLS regressions of olfactory ratio (olfactory bulb size versus cerebrum size) for palaeognaths (solid) and neognaths (dashed). Neognath data from Bang and Cobb [22]. Lithornis plebius data from Zelenitsky et al. [10]. Colour legend: orange, elephant birds; red, kiwi; yellow, cassowary; pink, emu; purple, moa; cyan, tinamous; green, rhea; tan, ostrich; blue, lithornithid; white, neognaths.