Table 2.

Estimates of the distribution of fitness effects of new mutations at zerofold nonsynonymous sites, intronic sites, 5′ UTR sites, 3′ UTR sites, and intergenic sites falling in different N_es ranges, and proportion of divergence driven to fixation by positive selection (α) and the rate of adaptive substitution relative to neutral divergence (ω) in P. tremula and P. tremuloides

		Percentage of mutations in Nes range
Species	Category	0–1	1–10	10–100	>100	α	ω
P. tremula	Zerofold	0.275 (0.263–0.277)	0.118 (0.115–0.132)	0.167 (0.163–0.196)	0.440 (0.408–0.447)	0.298 (0.289–0.335)	0.107 (0.103–0.121)
	3′ UTR	0.822 (0.807–0.829)	0.096 (0.096–0.119)	0.073 (0.067–0.077)	0.009 (0.003–0.011)	0.125 (0.115–0.151)	0.114 (0.104–0.137)
	5′ UTR	0.646 (0.614–0.653)	0.163 (0.148–0.232)	0.157 (0.150–0.161)	0.034 (0.005–0.050)	0.311 (0.299–0.365)	0.275 (0.263–0.324)
	Intronic	0.727 (0.722–0.731)	0.088 (0.088–0.089)	0.096 (0.096–0.096)	0.089 (0.085–0.095)	0.159 (0.151–0.167)	0.133 (0.125–0.141)
	Intergenic	1.000 (1.000–1.000)	0.000 (0.000–0.000)	0.000 (0.000–0.000)	0.000 (0.000–0.000)	0.215 (0.210–0.220)	0.273 (0.265–0.282)
P. tremuloides	Zerofold	0.264 (0.251–0.266)	0.131 (0.128–0.147)	0.194 (0.190–0.227)	0.411 (0.376–0.416)	0.338 (0.330–0.374)	0.122 (0.118–0.135)
	3′ UTR	0.853 (0.834–0.858)	0.095 (0.094–0.120)	0.051 (0.046–0.056)	0.001 (0.000–0.002)	0.094 (0.084–0.132)	0.086 (0.076–0.120)
	5′ UTR	0.632 (0.601–0.643)	0.225 (0.207–0.278)	0.140 (0.123–0.147)	0.004 (0.000–0.007)	0.345 (0.329–0.385)	0.305 (0.289–0.342)
	Intronic	0.748 (0.713–0.752)	0.091 (0.090–0.140)	0.096 (0.095–0.124)	0.065 (0.023–0.069)	0.140 (0.133–0.197)	0.118 (0.111–0.165)
	Intergenic	1.000 (1.000–1.000)	0.000 (0.000–0.000)	0.000 (0.000–0.000)	0.000 (0.000–0.000)	0.207 (0.200–0.213)	0.261 (0.250–0.271)

Ninety-five percent bootstrap confidence intervals are shown in parentheses.