Comparison of root radial hydraulic properties measured and simulated for various cell-scale hydraulic properties along a root. The three simulated distances from the tip correspond to different levels of apoplastic barrier development: endodermal Casparian strip (5 cm), suberized endodermis with passage cells (25 cm), and suberized endodermis and exodermal Casparian strip (50 cm). A, Comparison between experimental kr ranges in maize primary roots (colored rectangles and dashed line) and kr simulated from the cell scale in the maize hydraulic anatomy (consistent symbols across legends with impermeable and leaky apoplastic barriers in black and gray, respectively). B, Comparison between experimental mannitol σr ranges in maize principal roots (colored rectangles) and root segment σr arising from the cell scale in the maize hydraulic anatomy (σr values lower than 1 were obtained with the water-permeable apoplastic barrier).