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. 2018 Jul 5;68(1):168–181. doi: 10.1093/sysbio/syy051

Table 1.

Rate matrix for Markov chain describing transitions between states in multispecies coalescent with migration model with two populations (Inline graphic and Inline graphic) and three sequences (Inline graphic, Inline graphic, and Inline graphic)

  Inline graphic Inline graphic Inline graphic Inline graphic Inline graphic Inline graphic Inline graphic Inline graphic Inline graphic Inline graphic Inline graphic Inline graphic Inline graphic Inline graphic Inline graphic Inline graphic Inline graphic Inline graphic Inline graphic Inline graphic Inline graphic
Inline graphic Inline graphic Inline graphic Inline graphic   Inline graphic       Inline graphic Inline graphic Inline graphic                    
Inline graphic Inline graphic Inline graphic   Inline graphic   Inline graphic                           Inline graphic  
Inline graphic Inline graphic   Inline graphic Inline graphic     Inline graphic                       Inline graphic    
Inline graphic   Inline graphic Inline graphic Inline graphic       Inline graphic             Inline graphic            
Inline graphic Inline graphic       Inline graphic Inline graphic Inline graphic                     Inline graphic      
Inline graphic   Inline graphic     Inline graphic Inline graphic   Inline graphic               Inline graphic          
Inline graphic     Inline graphic   Inline graphic   Inline graphic Inline graphic                 Inline graphic        
Inline graphic       Inline graphic   Inline graphic Inline graphic Inline graphic       Inline graphic Inline graphic Inline graphic              
Inline graphic                 Inline graphic           Inline graphic     Inline graphic     Inline graphic
Inline graphic                   Inline graphic           Inline graphic     Inline graphic   Inline graphic
Inline graphic                     Inline graphic           Inline graphic     Inline graphic Inline graphic
Inline graphic                       Inline graphic     Inline graphic     Inline graphic     Inline graphic
Inline graphic                         Inline graphic     Inline graphic     Inline graphic   Inline graphic
Inline graphic                           Inline graphic     Inline graphic     Inline graphic Inline graphic
Inline graphic                 Inline graphic     Inline graphic     Inline graphic            
Inline graphic                   Inline graphic     Inline graphic     Inline graphic          
Inline graphic                     Inline graphic     Inline graphic     Inline graphic        
Inline graphic                 Inline graphic     Inline graphic           Inline graphic      
Inline graphic                   Inline graphic     Inline graphic           Inline graphic    
Inline graphic                     Inline graphic     Inline graphic           Inline graphic  
Inline graphic                                         Inline graphic

Note: Inline graphic and Inline graphic are mutation-scaled migration rates, and Inline graphic and Inline graphic are the coalescent rates. The state of the chain is given by the population IDs (Inline graphic or Inline graphic) and sequence IDs (such as Inline graphic, Inline graphic, Inline graphic). For example the initial state Inline graphic means that the three sequences Inline graphic, and Inline graphic are from populations Inline graphic, Inline graphic, and Inline graphic, respectively. States with three sequences are abbreviated, with the three sequences assumed to be in the order Inline graphic so that the sequence IDs are suppressed. Thus Inline graphic is ‘Inline graphic’. State Inline graphic means that two sequences remain in the sample, with the ancestor of sequences Inline graphic and Inline graphic is in population Inline graphic while sequence Inline graphic is in population Inline graphic. This is abbreviated ‘Inline graphic’, with the sequence ID ‘Inline graphic’ suppressed. ‘Inline graphic’ is an absorbing state in which only one sequence remains in the sample, in either Inline graphic or Inline graphic, after two coalescent events have occurred.