The TIGR brush. (A) Slice of a domestic cat tongue illuminated using UV dye and a black light. (B) A 3D-printed mimic displaying flexibility similar to the cat tongue. (C and D) Artificial grooming data for the 3D-printed mimic (C) and a human hairbrush (D). Each color represents a successive trial in rainbow order (red, orange, yellow, green, blue, purple, black). Force peaks indicate an encountered tangle. (E) Schematic displaying a relaxed, streamlined papilla encountering a tangle (Left), catching on a tangle (Center), and rotating until perpendicular to the tongue (Right).