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Rates of melanosome shape evolution and variation in metabolic rate across amniotes. Colours of tips correspond to mass-derived metabolic rate (dark blue: low, yellow: high) with fossils in grey. Darker branch colours indicate faster rates of melanosome evolution. Rate shifts are shown as filled circles (red: speed-up, blue: slow-down). Inset shows multivariate phylogenetic contrasts for melanosome morphology versus ancestral metabolic rates (r = 0.19, prand = 0.006).
