Enzymatic transfer of
α-PD-L1 to OT-1 T cells for enhanced
T-cell activation and specific killing: (A) Schematic illustration
of the interaction between OT-1 T cells and B16-OVA melanoma cells.
MHC molecules on B16-OVA present OVA antigen to OVA-specific TCR on
OT-1 T cells to induce activation, while PD-L1 on B16-OVA interact
with PD-1 on OT-1 T cells to inhibit the T-cell effector function.
(B) Schematic illustration of the blockade of PD-1–PD-L1 pathway
via α-PD-L1 conjugated on the surfaces of OT-1 T cells. The in situ blockade could enhance T-cell activation and the
killing of cancer cells. (C) Analysis of PD-L1 antigen binding on
OT-1 T cells for different treatments. Mean ± SD (error bars).
(D) Analysis of PD-L1 antigen binding of OT-1 T cells modified with
various concentrations of GF-α-PD-L1. Mean ± SD (error
bars). (E) Quantifying α-PD-L1-OT-1 T cell conjugates
mediated killing of B16-OVA cells. Mean ± SD (error bars), representative
graph from three independent experiments. OT-1 T cells conjugated
with rIgG and P14 T cells conjugated with α-PD-L1 were shown
as negative control. (F) Comparison of OT-1 T cells and α-PD-L1-OT-1
T cell conjugates in killing B16-OVA at different effector-to-target
cell ratios. Mean ± SD (error bars). (G) Killing activities of
OT-1 T cells and α-PD-L1-OT-1 T cell conjugates constructed
at different GF-α-PD-L1 concentrations. Mean ± SD (error
bars). (H) Comparison of α-PD-L1-OT-1 T cell conjugates with
OT-1 T cells with or without additionally added free α-PD-L1
(5 μg/mL) in killing B16-OVA (3 h incubation). (I) IFN-γ
secretion analysis of modified or unmodified OT-1 T cells in a B16-OVA
cell co-culture assay. P14 T cells conjugated with α-PD-L1 were
used as a negative control. Mean ± SD (error bars), representative
graph from three independent experiments. (J) Microscopy images of
OT-1 T cell-induced killing of B16-OVA with or without α-PD-L1
labeling. The blue arrow indicates fewer cancer cells, and the purple
arrow indicates larger clusters of T cells. Scale bar: 50 μm.
(K) Analysis of OT-1 T cell proliferation via CFSE dilution in a B16-OVA
cell co-culture assay. Mean ± SD (error bars). In all figures,
ns, P > 0.05; **P < 0.01;
***P < 0.001; ****P < 0.0001;
one-way
ANOVA followed by Tukey’s multiple comparisons test, two-way
ANOVA followed by Sidak’s multiple comparisons test.