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. 2019 Jan 2;14(1):e0208694. doi: 10.1371/journal.pone.0208694

Table 1. Prey items of subadult and adult grey seal males indicated by gut morphological i.e. hard part (HP, mass %) and DNA (sequence %) prey proportions, and the frequencies of occurrence (Freq %).

Prey items Subadult males Adult males
HP (%)
n = 37
DNA (%)
n = 42
HP (%)
n = 51
DNA (%)
n = 44
Prop Freq Prop Freq Prop Freq Prop Freq
Baltic herring Clupea harengus 46.7 59.5 39.8 69.0 29.0 58.8 24.6 63.6
Perch Perca fluviatilis 13.6 21.6 11.0 21.4 13.0 35.3 11.1 36.4
Eelpout Zoarces viviparus 10.3 13.5 6.1 28.6 7.0 17.6 8.5 18.2
Bream Abramis brama ND ND 6.0 16.7 3.4 5.9 10.8 22.7
Roach Rutilus rutilus 3.0 8.1 3.4 11.9 5.0 13.7 5.1 31.8
Sprat Sprattus sprattus 2.6 13.5 9.2 23.8 0.5 7.8 2.9 22.7
Pikeperch Sander lucioperca 4.3 5.4 2.4 16.7 1.5 7.8 5.8 13.6
Common whitefish C. lavaretus 0.3 2.7 ND ND 9.2 21.6 ND ND
Atlantic salmon Salmo salar ND ND 0.6 9.5 1.5 3.9 6.2 6.8
European eel Anguilla anguilla 3.0 5.4 1.7 7.1 1.6 3.9 1.0 6.8
Three-spined stickleback G. aculeatus < 0.1 2.7 5.3 7.1 ND ND <0.1 2.3
Cod Gadus morhua 0.2 2.7 3.3 11.9 1.1 5.9 0.3 2.3
Burbot Lota lota ND ND 3.4 9.5 0.5 2.0 0.5 4.5
Pike Esox lucius ND ND ND ND 1.8 2.0 2.2 6.8
Isopod crustacean Saduria entomon 2.9 8.1 ND ND <0.1 3.9 ND ND
Sea trout Salmo trutta 0.9 2.7 ND ND <0.1 2.0 1.2 2.3
Ruffe Gymnocephalus cernua 1.0 5.4 0.3 4.8 <0.1 2.0 0.8 2.3
European flounder Platichthys flesus ND ND ND ND 1.7 5.9 ND ND
Turbot Scophthalmus maximus ND ND ND ND 1.4 2.0 ND ND
Four-horned sculpin M. quadricornis 0.4 2.7 ND ND 0.9 5.9 ND ND
Smelt Osmerus eperlanus 0.5 8.1 < 0.1 2.4 0.3 5.9 0.2 6.8
Sand goby Pomatoschistus minutus ND ND 0.7 2.4 ND ND < 0.1 4.5
Common dab Limanda limanda ND ND ND ND 0.7 2.0 ND ND
White bream Blicca bjoerkna ND ND < 0.1 4.8 ND ND 0.2 4.5
Tench Tinca tinca ND ND ND ND 0.1 2.0 ND ND
Black goby Gobius niger ND ND < 0.1 2.4 <0.1 2.0 ND ND
Rainbow trout Oncorhynchus mykiss ND ND < 0.1 2.4 ND ND ND ND
Cyprinids Cyprinidae 5.0 16.2 ND ND 14.4 31.4 ND ND
Whitefishes Coregonus spp ND ND 2.2 11.9 0.3 3.9 13.2 27.3
Percids Percidae 2.2 5.4 ND ND 3.1 11.8 ND ND
Sculpins Cottidae < 0.1 2.7 2.4 4.8 <0.1 3.9 1.8 13.6
European flounder or plaice
Plat. flesus or Pleur. platessa
ND ND < 0.1 2.4 ND ND 2.4 4.5
Clupeids Clupeidae 0.3 2.7 ND ND 2.0 5.9 ND ND
Daces Leuciscus sp. ND ND 1.2 2.4 ND ND 1.0 2.3
Sand lances Ammodytidae 1.1 8.1 0.7 4.8 ND ND ND ND
Lampreys Petromyzontidae 1.7 2.7 ND ND ND ND ND ND
Gobies Gobiidae < 0.1 5.4 ND ND <0.1 15.7 ND ND

The items identified at species levels were listed first (in the order of decreasing average proportion in the subadult and adult diets, indicated by the HP and DNA analyses) and those identified at species group levels were listed after the species. The number of subadult and adult males from which both HP and DNA were recorded was 33 and 40, respectively.