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Map of the region of the S. pyogenes chromosome that includes the has operon encoding enzymes required for hyaluronic acid synthesis. The other genes shown appear not to be involved in capsular polysaccharide synthesis or surface expression. In some strains insertion sequence IS1239′ is present approximately 50 nucleotides upstream of the has operon promoter (adapted from the genome sequence of M1 strain SF370, GenBank accession number AE004092 [93]).
