Fig. 5.

Ancestral area reconstruction based on BEAST trees constrained using nuclear species tree (HCX(Cu,Ju)) topology and the DIVALIKE +J model in BioGeoBEARS, as implemented in RASP 4.0. The inset shows the division of the distribution area of the five genera into four operational areas (North America, Africa, Asia and Europe). The pie at each node represents the reconstructed ancestral area; different colours of circular sectors in a pie represent the relative probabilities of different ancestral areas at a node.