Average Shannon diversity varies according to time of sampling. Shannon diversity at 1230 h was significantly higher than that at every other time point, excluding 1600 h; diversity at 2000 h was significantly lower than that at every other time point, excluding 1100 h (Kruskal Wallis pairwise, corrected P value of <0.05; n = 17 fish at 1100 and 2000 h and 18 fish at 1230, 1400, and 1600 h).