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. 2019 Jan 29;9:825. doi: 10.1038/s41598-018-36069-w

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© The Author(s) 2019

Open Access This article is licensed under a Creative Commons Attribution 4.0 International License, which permits use, sharing, adaptation, distribution and reproduction in any medium or format, as long as you give appropriate credit to the original author(s) and the source, provide a link to the Creative Commons license, and indicate if changes were made. The images or other third party material in this article are included in the article’s Creative Commons license, unless indicated otherwise in a credit line to the material. If material is not included in the article’s Creative Commons license and your intended use is not permitted by statutory regulation or exceeds the permitted use, you will need to obtain permission directly from the copyright holder. To view a copy of this license, visit http://creativecommons.org/licenses/by/4.0/.

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Crystal structure of NL BdGUS. (a) Monomeric tertiary structure and dimeric quaternary structure of BdGUS (orange). A single monomer is highlighted in orange and the identical monomer is shown with the glycosyl hydrolase 2 (GH2) in white, and the C-terminal domains (domain of unknown function, DUF; carbohydrate binding motif; CBM) in red and blue, respectively. (b) Active site of BdGUS with the CBM loop indicated by an arrow. Catalytic glutamates are boxed.