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. 2019 Jan 29;9:847. doi: 10.1038/s41598-018-38155-5

Table 4.

Characterization of biological and ecological traits and their states used for beetle species assignment for purposes of this study.

Trait State Description No of assigned species
trophic guild predators species which kill and feed on other animals 91
herbivores species which feed on living (green) plant tissues 101
mycetophages species which feed on living mushrooms 17
saprophages species which feed on dead organisms (except dead wood) 35
cambioxylophages species which feed on wood and phloem 53
habitat type forest-dwellers species which live only in forested habitats 112
open land-dwellers species which live only in grassland habitats 131
ubiquitous species which could live in all types of terrestrial habitats 54
microhabitats coprophilous species which inhabit animal feces 20
epigeic species which inhabit soil surface (free living) 38
mycetophilous species which inhabit mushrooms 15
herbophilous species which inhabit living plants 128
hygrophilous species which inhabit wet habitats (but not aquatic) 25
necrophilous species which inhabit carcases 5
cambioxylophilous species which inhabit inside trees 66
distribution continental species of xeric habitats in Pontic and Pannonian regions (mostly steppes) 25
mediterranean species of dry habitats in Mediterranean basin (the Balkans) 17
mountainous species of high mountain zones (the Sudetes, the Carpathians and the Balkan Mts.) 11
temperate species widespread in central Europe except above listed areas 244
range fragmented species with fragmented and isolated populations 36
widespread species with continuous ranges 261
thermal preferences neutral species which wide tolerance to temperatures 250
warm species which prefer only warm temperatures 47
reproduction mode bisexual species which reproduce only bisexually 292
parthenogenetic species which parthenogenetic reproduction (at least in central Europe) 5
body size large species larger than 20 mm length 23
medium species of 10–20 mm length 174
small species smaller than 10 mm length 100
mobility mobile species winged and capable to fly 274
sedentary species apterous (flightless) 23