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. 2019 Jan 30;9:973. doi: 10.1038/s41598-018-37801-2

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© The Author(s) 2019

Open Access This article is licensed under a Creative Commons Attribution 4.0 International License, which permits use, sharing, adaptation, distribution and reproduction in any medium or format, as long as you give appropriate credit to the original author(s) and the source, provide a link to the Creative Commons license, and indicate if changes were made. The images or other third party material in this article are included in the article’s Creative Commons license, unless indicated otherwise in a credit line to the material. If material is not included in the article’s Creative Commons license and your intended use is not permitted by statutory regulation or exceeds the permitted use, you will need to obtain permission directly from the copyright holder. To view a copy of this license, visit http://creativecommons.org/licenses/by/4.0/.

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Radiographic findings in a 3-month-old affected dog. (a) The oldest affected dog (litter 1) had a crouched, planti- and palmigrade stance and gait. (b) Laterolateral radiograph of the head. Both maxilla and mandible are shortened and the lower jaw is malformed with an angulated mandibular body and an elongated mandibular ramus (arrowhead). (c) Laterolateral radiograph of the lumbar spine. The vertebral bodies are short, the end plates are poorly mineralized and the dorsal borders of the vertebral spinous processes are irregular. Mediolateral radiograph of (d) the right front and (e) right hind limbs. The long bones are hourglass-shaped (open arrowheads) and have hypoplastic marrow cavity, as well as hypoplastic cortical bone. Carpal bones and distal femoral epiphyses show irregular mineralization (arrowheads).