Figure 1.

(a) measurement of drone heads (H) was perpendicular to the body axis at the widest point of the eyes and measurement of thorax width was measured as the distance between the tips of the two tegulae (T); (b) length of the dissected forewings (W) was taken as the length from the base of the costa to the tip of the wing, crossing through the intersection of the 2nd and 3rd sub-marginal cells with the 2nd marginal cell. The average of both wings was analyzed; (c) length of each mucus gland (MG) along the central axis was measured from the distal bulb to the ejaculatory duct. Lengths were averaged for analysis. Length of each seminal vesicle (SV) was measured along the central axis from the base of the vas deferens to the mucus gland and an average of both was used for analysis.