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. 2019 Feb 20;9:2347. doi: 10.1038/s41598-019-38818-x

Table 1.

The frequency and expressions of key dental traits in Dushan 1 and comparative samples.

Dushan 1 East Asia
archaic Homo
East Asia
EMH
East Asia
RMH
Java
H. erectus
Neanderthal Global
H. sapiens
Trait comparisons (Frequency and specimen number)
3 I1 Labial convexity 1/2
(EP-2)
92.8% (13/14) 42.9% (3/7) 9.6% (8/83) 100%
N = 21
30.8%
N = 91
6 I2 Interruption groove
present
1/2
(Hs)
0% (0/7) 0% (0/6) 43.6% (92/211) 10.4–65.0%
N = 5693
9 P3 Transverse crest present 0/2
(Hs)
17.7% (3/17) 20% (1/5) 4.2% (3/71) 33.3% (3/9) 6.7% N = 15 2% (6/283)
10 P4 Transverse crest present 0/2
(Hs)
21.4% (3/14) 25% (1/4) 2.9% (2/70) 50% (4/8) 12.4% N = 16 2% (5/279)
13 P3 Bifurcated essential crest 2/2
(EP-2)
100% (15/15) 0% (0/4) 44.2% (23/52) 53.3% N = 15 11.1% N = 126
14 P4 Bifurcated essential crest 2/2
(EP-2)
100% (16/16) 55.3% (26/47) 78.6% N = 14 9.4% N = 106
17 P3 Mesial or distal accessory ridge 2/2
(Hs)
93.3% (14/15) 50% (2/4) 43.3% (26/60) 36.4% N = 11 23.4–57.0%
N = 749
18 P4 Mesial or distal accessory ridge 2/2
(Hs)
93.8% (15/16) 67.6% (46/68) 41.7% N = 12 33.0–80.2%
N = 749
19 P3 Mesial or distal accessory ridge 2/2
(Hs)
100% (10/10) 25% (1/4) 52.0% (40/77) 90% N = 20 11.9% N = 13521
20 P4 Mesial or distal accessory ridge 2/2
(Hs)
100% (8/8) 33.3% (1/3) 46.6% (34/73) 70% N = 20 23% N = 115
21 P3 Accessory marginal tubercle 2/2
(EP-2)
93.3% (14/15) 25% (1/4) 19.2% (10/52)
22 P4 Accessory marginal tubercle 2/2
(EP-2)
93.8% (15/16) 14.9% (7/47)
24 P4 Two-rooted premolars 2/2
(EP-2)
40% (2/5) 0% (0/6) 4.3% (2/47)
27 Three-rooted P3 2/2
(EP-1)
0% (0/9) 0% (0/5) 0% (0/46) 0% (0/599)
28 P3 Tomes’s Root 2/2
(EP-2)
64.8% (142/219) 19.9% N = 1371
39 M1 Carabelli’s trait
(Dushan 1 has special expression)
2/2
(EP-1+ EP-2)
45% (9/20) 21.4% (3/14) 9.7% (27/279) 100% (2/2) 50%
N = 20
1.9–36.0%
N = 2426
49 M1 protostylid
(Dushan 1 has special expression)
2/2
(EP-1+ EP-2)
53.9% (7/13) 18.2% (2/11) 18.3% (63/345) 3.2%
N = 31
18.7%
N = 2362
60 Three-rooted M1 2/2
(Hs)
7.7% (1/13) 0% (0/15) 24.2% (15/62) 0–31.1%
62 Complex occlusal morphology at EDJ of molars 5/5
(EP-2)
100% (16/16) 15.2% (5/33) 5.2% (8/153)
63 Buccal basal swelling
C1- M3
C1- M3
2/2
2/2
(EP-2)
54.6–63.6%
N = 19
55.6–88.9%
N = 22
14.6%
N = 82
12.9%
N = 70
0–25.7%
N = 101
0–12.9%
N = 126
65 Crown buccal vertical groove complex Present
(Unique + EP-1)
Absent Absent Absent
N = 5
Weakly expressed
(La Quina H18)
66 Tooth size proportions
I1-C1/P3-M3
0.32
(Hs)
0.40 0.37 0.37 0.42 0.36
67 Tooth size proportions
I1-C1/P3-M3
0.26
(Hs)
0.27 0.25 0.26 0.31 0.26
68 Molar size % increase
M1– M2
−0.5/+2.6 N = 2
(EP-2)
Mean = −1.4
− 11.5/+6.3 N = 6
Mean = 1.2
−3.1/+9.9  N = 8
Mean = −5.0
−14.2/+4.8 N = 174
+4/+9 N = 5 −15/+5 N  = 250
69 Molar size % increase
M2– M3
−1.3/−0.9 N = 2
(EP-2)
Mean = −2.8
−13.1/+6.3 N = 5
Mean = −3.7
−6.4/+4.7 N = 5
Mean = −2.6
−18.9/+18.0 N = 124
−10/0 N = 5 −15/+14 N = 211

Morphological status of Dushan Cave in the parentheses.

‘EP-1’: a primitive condition shared with the Africa Austrapithecus and early Homo.

‘EP-2’: a primitive condition shared with the Eurasia archaic Homo.

‘Hs’: a derived condition shared with H. sapiens.

‘Hs-Eu’: a derived condition shared with European H. sapiens or Neanderthal.

‘unique’: a unique condition restricted to Dushan Cave.