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The novel aggrecanase is inhibited by tissue inhibitor of metalloproteinase (TIMP)-3. Western blot analyses show that TIMP-3 inhibits retinoic acid-induced aggrecanolysis at FREEE 1467↓1468 GLGSV (a) and DIPEN 341↓342 FFGVG (b) in TS-4/5Δcat cartilage explants.
