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. 2019 Feb 27;12:42. doi: 10.1186/s13068-019-1384-0

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© The Author(s) 2019

Open AccessThis article is distributed under the terms of the Creative Commons Attribution 4.0 International License (http://creativecommons.org/licenses/by/4.0/), which permits unrestricted use, distribution, and reproduction in any medium, provided you give appropriate credit to the original author(s) and the source, provide a link to the Creative Commons license, and indicate if changes were made. The Creative Commons Public Domain Dedication waiver (http://creativecommons.org/publicdomain/zero/1.0/) applies to the data made available in this article, unless otherwise stated.

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Fig. 9 — Proposed conversion of cellulose to glucuronic acid-containing cello-oligosaccharides by HiPMO1. HiPMO1 oxidizes the primary alcohol group at C-6 of cellulose to the aldehyde group and the carboxylic group (ROH → RCHO → RCOOH). The resulting glucuronic acid-containing cello-oligosaccharides might be further cleaved via hydrolysis by beta-glucuronidase (GUS) and beta-glucosidase (BGL) to yield glucose, glucuronic acid, and saccharic acid, and cleaved via beta-elimination by PL to produce unsaturated cello-oligosaccharides