PILS6 is a negative regulator of organ growth. (A–C) PILS6 affects organ growth. Scanned images and measurements show that total root length (A), cotyledon area (B), and rosette area (C) are affected in the pils6 mutants and PILS6OE compared with the wild-type control. (D) PILS6 affects cell division. GUS images and measurements of CYCB1;1::GUS expression (first leaf in Upper images, root in Lower images) and root meristem distribution show that cell division is affected in the pils6 mutant and PILS6OE compared with the wild-type control. The black dashed boxes represent the ROIs used to quantify CYCB1;1::GUS signal intensity. The red dashed line shows how the length of CYCB1;1::GUS distribution in the meristem was quantified. [Scale bars, 500 μm (A–C) and 100 μm (D).] n = 49–55 roots (A), 50–56 cotyledons (B), 28 rosettes (C), 10–12 first leaves (D), and 28 roots (D). *P = 0.01–0.05, **P = 0.001–0.01, ***P < 0.001, one-way ANOVA.