Phylogenies of selected repair and nucleotide metabolism genes of the pithovirus-iridovirus-marseillevirus group that includes Loki’s Castle viruses. (A) SbcCD nuclease, ATPase subunit SbcC. (B) SbcCD nuclease, nuclease subunit SbcD. (C) Exonuclease V. (D) DNMP kinase. The numbers at the internal branches indicate (percent) local likelihood-based support. GenBank protein identifiers (IDs), wherever available, are shown after each “@” symbol. Taxon abbreviations are as follows: A, Archaea; B, Bacteria; E, Eukaryotes; N, NCLDV; DP, DPANN group; TA, Thaumarchaeota; Ea, Euryarchaeota; FC, Bacteroidetes; Fu, Fusobacteria; Pr, Proteobacteria; Te, Firmicutes; un, unclassified Bacteria; Op, Opisthokonta; Pi, “Pithoviridae”; Ac, Ascoviridae; As, Asfarviridae; Ma, Marseilleviridae; Mi, Mimiviridae; Pa, Pandoraviridae; Ph, Phycodnaviridae; V ds, double-strand DNA viruses.