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. 2019 Mar 11;18:52. doi: 10.1186/s12934-019-1102-4

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© The Author(s) 2019

Open AccessThis article is distributed under the terms of the Creative Commons Attribution 4.0 International License (http://creativecommons.org/licenses/by/4.0/), which permits unrestricted use, distribution, and reproduction in any medium, provided you give appropriate credit to the original author(s) and the source, provide a link to the Creative Commons license, and indicate if changes were made. The Creative Commons Public Domain Dedication waiver (http://creativecommons.org/publicdomain/zero/1.0/) applies to the data made available in this article, unless otherwise stated.

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Fig. 1 — The workflow of using SCRaMbLE to improve and analyze alkali tolerance in yeast. Three sections in varied colors indicated three sequential steps of the whole process including construction of inducible SCRaMbLE system, screening for specific phenotype, correlation analysis of phenotype and genotype