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. 2019 Mar 11;14(3):e0213031. doi: 10.1371/journal.pone.0213031

Table 1. Slipping behaviour ethogram.

Ethogram of collective fish behaviour during escape from purse seines.

Behavioural unit Type Description Hypothesised welfare impact
No escape State Discharge channel (or its immediate area) in field of view of cameras, but with no fish escaping the net Uncertain–positive impacts resulting from maintenance of schooling; negative impacts resulting from continued risk of impact of capture stressors
Single fish Event Discharge channel (or its immediate area) in field of view of camera, with a single fish escaping the net Moderate–loss of schooling advantages in an obligatory schooling species
Small group Event Discharge channel (or its immediate area) in field of view of camera, with a small group1 of fish (<30) escaping the net with a coordinated schooling structure Minor–school structure remains intact, low probability of contact related injuries
Orderly State Discharge channel (or its immediate area) in field of view of camera, with a large group of fish (>100) escaping3 the net with a coordinated schooling structure Minor–school structure remains intact, low probability of contact related injuries
Disorderly State Discharge channel (or its immediate area) in field of view of camera, with a large group of fish (>100) escaping3 the net without a coordinated schooling structure, often with collisions between fish and into gear High–schooling structure lost in an obligatory schooling species, high probability of contact related injuries
Disagreement State Discharge channel (or its immediate area) in field of view, but with disorderly behaviour and orderly behaviour evident on different cameras Minor / High–as for orderly or disorderly above.
Return Event Discharge channel (or its immediate area) in field of view of camera and an escape taking place, with ≥ 1 fish of the group re-entering the net Uncertain–positive impacts resulting from joining large school inside the net; negative impacts resulting from increased risk of impact from capture stressors
Not in view State Either: 1) both camera views obscured, or 2) discharge channel (or its immediate area) not in field of view of either camera NA

1A group was defined as >1 fish in close proximity (less than approximately 5 body lengths distance between individuals)

2A coordinated structure was defined as a polarized group of fish, capable of collective schooling movements

3The start and end of an escape for groups of >100 fish was defined as when the first and last fish passed the discharge channel threshold.