Optimal conditions for nitrification are represented, including low light, the presence of oxygen and high ammonium (NH4+) concentrations. By controlling the availability of N forms (mainly NH4+ and nitrate–NO3-), nitrification influences phytoplankton assimilation and community structure, as well as facilitating permanent N removal through coupled nitrification-denitrification. Ammonia oxidation (AO) can be conducted by both ammonia-oxiding archaea (AOA) and ammonia-oxidizing bacteria (AOB), whereas nitrite oxidation is conducted by nitrite-oxidizing bacteria (NOB). AO also produces nitrous oxide (N2O) as a by-product, a potent greenhouse gas.