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Fe-atom binding site. (a) shows the Fe atom bound by the protein through a trigonal bipyramidal coordination, with Asp193, His260 and C5N of NADP at the corners of the trigonal plane and His197 and His272 in the axial positions. The NADP is in van der Waals contact with the Fe atom, which is presumably in the ferric state. The view in (b) is rotated approximately 90° from that in (a). Here it can be seen that the nicotinamide ring is edge-on, with C5N directed towards the iron.
