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The maximum likelihood phylogeny categorized Lon homologs in four major clades. Clades I and II contain Lon of prokaryotic origin diverged in Archaea and bacteria, respectively. Clade III includes the peroxisomal Lon homologs, whereas Clade IV contains the bona fide mitochondrial Lon homologs of fungi, animals, and algae, and Lon homologs from land plants dual-targeted to mitochondria and chloroplasts.
