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Components of an SEIR model as applied to environmental transmission during a single outbreak season (t). Susceptible hosts (S) move across the landscape (A) and contact infectious LIZs (I) and become exposed (E; B). As they leave the LIZ, they may succumb to infection and die, becoming a new LIZ, establishing in time t and persisting across future time periods (t + 1,…,n) (C) or recover (R) and survive to a future time period (D).
