A. Schematic of CC2D1A as a signaling scaffold for PDE4D and PKA upstream of CREB. B-E. Conditional removal of Cc2d1a in cKO male mice causes increased phosphorylation and activity of PDE4D (B.), leading to reduced levels of cAMP (C.), and decreased phosphorylation of both PKA (D.) and CREB (E.) in hippocampal lysates. F-I. No changes in PDE4D (F.), cAMP (G.), PKA (H.) or CREB (I.) are found in the female hippocampus. Results expressed as mean ± SEM. Data averages and statistical information are reported in the Results. Two-tailed t-test with equal variance * p<0.05, ** p<0.01, *** p<0.001