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. 2019 Apr 24;13:22. doi: 10.3389/fncir.2019.00022

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Copyright © 2019 Lewis, Purdy, Ahmad and Hawkins.

This is an open-access article distributed under the terms of the Creative Commons Attribution License (CC BY). The use, distribution or reproduction in other forums is permitted, provided the original author(s) and the copyright owner(s) are credited and that the original publication in this journal is cited, in accordance with accepted academic practice. No use, distribution or reproduction is permitted which does not comply with these terms.

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A diagram of the network with arrows indicating the main connections. (1) Motor input shifts the activity in the location layer, which consists of a set of independent grid cell modules. (2) The active location cells provide modulatory predictive input to the sensory layer. (3) Sensory input activates cells in the sensory layer. (4) The location is updated by the new sensory representation.