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. 2019 May 2;9:6856. doi: 10.1038/s41598-019-43313-4

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© The Author(s) 2019

Open Access This article is licensed under a Creative Commons Attribution 4.0 International License, which permits use, sharing, adaptation, distribution and reproduction in any medium or format, as long as you give appropriate credit to the original author(s) and the source, provide a link to the Creative Commons license, and indicate if changes were made. The images or other third party material in this article are included in the article’s Creative Commons license, unless indicated otherwise in a credit line to the material. If material is not included in the article’s Creative Commons license and your intended use is not permitted by statutory regulation or exceeds the permitted use, you will need to obtain permission directly from the copyright holder. To view a copy of this license, visit http://creativecommons.org/licenses/by/4.0/.

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Eleven encounter types seen in simulations and observed data sets (Figs. 3 and 4) characterised by the five behaviour indices Mean Maximum Aggression Index (MMAI), Mean Behaviour Index aggressive (MBI_agg), Aggression Index (AI), Mean Behaviour Index peaceful (MBI_pcf), and Mean Maximum Peaceful Index (MMPI). This non-exhaustive collection of encounter types showcases that for characterising some encounter types (❶, ❷, ❿, ⓫; from simulations, not in observed data in this study), any index can be used but that for all others, the combination of multiple indices helps capturing behavioural complexity. Encounter type ❶ represents instant killing; ❷ killing after brief peaceful interactions; ❸ prolonged aggression; ❹ prolonged aggression alongside brief peaceful interactions; ❺ prolonged aggressive behaviour with brief peaceful behaviour; ❻ brief aggressive behaviour, brief peaceful behaviour, and prolonged neutral behaviour; ❼ brief aggression and prolonged peaceful behaviour; ❽ brief aggression with prolonged peaceful interactions; ❾ peaceful behaviour alongside ignoring; ❿ continuous trophallaxis alongside brief antennation; ⓫ trophallaxis throughout.