Flocs can be dispersed by cellulase or harsh vortexing. (A) Treatment of ZM4 flocs with cellulase, proteinase K, or DNase. Representative trends from one of two replicates are shown. (B) Effect of gentle (5 s at level 3) or harsh (10 pulses at level 10) vortexing on floc dispersal with (+) or without (–) a 3-h cellulase treatment. Error bars, standard deviations (n = 4). Asterisks indicate a significant difference (**, P < 0.01) by one-way analysis of variance with the Holm-Sidak posttest; ns, no significant difference.