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. 2019 Mar 4;212(1):245–265. doi: 10.1534/genetics.119.302000

Table A1. Summary of main results.

Landscape type Max allowed epistasis type Hardness of reaching local optima Proved in...
Smooth magnitude () Easy for all strong-selection weak-mutation (SSWM) dynamics Section B
Semismooth sign (Inline graphic,Inline graphic) Hard for SSWM with random fitter mutant Theorems 15,
or fittest mutant dynamics 20, and 24
Rugged reciprocal sign (Inline graphic) Hard for all SSWM dynamics: initial genotypes with all adaptive paths of exponential length Corollary 28
Hard for all evolutionary dynamics (if FP ≠ PLS) Theorem 27
Easy for finding approximate local peaks with moderate optimality gap: selection coefficients can drop-off as power law Theorem 33
Hard for approximate local peaks with small optimality gap: Theorem 35
selection coefficient cannot drop-off exponentially Corollary 36

Each landscape type (column 1) is characterized by the most complicated permitted type of epistasis (column 2; see A.1). Based on this, there are families of this landscape type that are easy or hard under progressively more general dynamics (column 3), which is proved in the corresponding part of the appendix (column 4).