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. 2019 Apr 9;47(9):4442–4448. doi: 10.1093/nar/gkz246
Category 1 (∈ rank) Category 2 (∉ rank)
Class 1 (∈ homoplasic trait) k n - k
Class 2 (∉ homoplasic trait) M - k N - M - n + k